Mill Run is an urban tributary to French Creek, one of the most biologically diverse streams in the Commonwealth of Pennsylvania, and home to more species of fish (89) and freshwater mussels (27) than anywhere in the state. Mill Run originates at the outfall of Tamarack Lake, a flood control and recreation reservoir located in West Mead Township. From there Mill Run flows through the city of Meadville to the confluence of French Creek. It is located in Crawford County, in Northwestern Pennsylvania.

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The Mill Run watershed covers 5,543 acres. Mill Run spans 3.75 miles in length, and has three primary functions today: it provides an aesthetic landscape in an urban community, supports biodiversity by providing habitat for many native species, and fosters community interaction at several locations throughout the City.